http://content.cdlib.org/view?docId=kt438n99wc;NAAN=13030&doc.view=frames&chunk.id=d0e442&toc.depth=1&toc.id=d0e442&brand=calisphere "
Spawning Season
The spawning season of trout varies to some extent with the locality and the temperature of the water. By nature, eastern brook and brown trout are fall spawners, while rainbows spawn in the spring (a complete list of the common and scientific names of the salmons and trouts of California is contained in the section The Classification of Fishes). In some California streams with runs of anadromous fishes, such as the Trinity River, there is hardly a month in the year when some strain of fish is not spawning. As an example, spring-spawning king salmon arrive in the upper river section at Lewiston the latter part of June. They are then followed by the summer and fall runs, which extend the salmon spawning season to mid-November. The king salmon runs overlap the silver salmon runs, which extend from November through January. The silvers in turn overlap the steelhead, which spawn during the period January through June. Actually, the steelhead of the Trinity River may be divided into those of the spring run (fish in general entering and migrating upstream on dropping stream levels while quite green, and spawning in the following season), and those of the fall run (fish in general entering on rising stream levels with sexual products in various stages of development, but spawning within the same season).
There are several ways in which the spawning season can be advanced. These include selective breeding, use of artificially controlled light, and injection of pituitary hormones. Selective breeding has been highly effective, and through this process the fall-spawning rainbow commonly referred to as the Hot Creek strain has been developed. Even though selective breeding is a very slow process, the results not being evident until several generations of offspring have been dealt with, the results are permanent and once the strain of fish has been developed it can be perpetuated by proper selection. It has been clearly demonstrated that the spawning time of trout can be advanced by the injection of pituitary hormones; however, this method is still in its infancy and, before the average hatcheryman will be benefited by its use, much needs to be learned regarding it. It offers interesting possibilities for crossbreeding varieties which spawn at different times of the year.
Effect of Temperature on Brood Stock and Eggs
Probably no other single thing regulates the development of eggs and growth of fish as much as temperature. It has been shown that rainbow trout reared in water held fairly constant at 60 degrees F. grow in length at the rate of about one inch per month. At 45 degrees F. they grow at less than one-quarter inch per month.
It is quite generally agreed that yearling and adult rainbow can withstand temperatures up to 78 degrees F. for short periods of time without harmful effect. It has also been shown that in order to produce eggs of good quality, rainbow spawners must be held at water temperatures not exceeding 56 degrees F., and preferably not above 54 degrees F., for a period of at least six months before spawning.
In order to get rainbow brood stock to grow rapidly and spawn when two years old, it is common practice to rear them the first 16 months of their life at a location where water temperatures are fairly constant at 60 degrees F., and then to transfer them to a location with a water temperature below 54 degrees F. to mature.
Just as water temperatures which are too warm (higher than 56 degrees) adversely affect egg development in rainbow and king salmon spawners, so do water temperatures which are too cold (42 degrees or lower) affect the development and incubation of trout and salmon eggs. In one experiment in which mature adult king salmon females, nearly ready to spawn, were placed in water ranging from 34 to 38 degrees, none of the females ripened and all died before spawning.
In an attempt to incubate king salmon eggs at a constant 35-degree temperature, mortality was practically 100 percent. Eggs held at water temperatures of 42.5 degrees or higher developed with only normal loss. Salmon eggs which had been held in water slightly above 42 degrees for a period of six days or longer could then tolerate colder temperatures without excessive mortalities. It is safe to say that the eggs of rainbow trout and king salmon will not develop normally in the fish if constant water temperatures above 56 degrees F. are encountered. It also follows that both rainbow trout and king salmon eggs cannot be incubated in water below 42 degrees F. without excessive loss."
Just to offer a sense of how variable all of this is, and how little of it is " written in stone," and how many factors there are that can influence when one or the majority of any population develop to any point.
Maybe the cohos are noticeable in years when there are low numbers of steelhead, and not so noticeable in years of high numbers of steelhead. There are a LOT of variables to try to juggle.